Two of the buttons on the left of the Nikon D4 remain the same, but the Command Lock button is replaced by the Metering mode button note the lack of a Metering Mode dial on the right of the pentaprism housing. A two-position collar switch selects between AF and MF, and the button in the middle works in conjunction with the Nikon D4's rear Command dial to set AF mode and AF area. The old CSM arrangement goes away in favor of the D7000's AF switch. Though they've moved a little, the monaural microphone, Self-timer lamp, Function button, Depth of Field preview button, and lens release buttons are in the same positions on the Nikon D4. Overall, the Nikon D4's body itself is more contoured, with fewer flat surfaces still, most items are placed similarly enough that seasoned D3S users will still feel at home. We won't go over every contour that's changed, because nearly every one has changed, at least on the front. Overall feel in the hand is similar to the D3S, but there are many cosmetic and ergonomic differences to the Nikon D4 worthy of note. Nearly the same size as the D3S, the Nikon D4 is only a fraction of an inch shorter, measuring 6.3 x 6.2 x 3.6 inches (160 x 156.5 x 90.5mm), and it weighs 47.3 ounces (2.95 pounds 1,340g) body-only with battery and XQD memory card. There's a whole lot more to the Nikon D4, which we'll cover in the Technical section below. Video is also upgraded from the D3S's 720p to 1080p at 30fps. The 51-point AF array improves sensitivity to -2 EV, and can accommodate lenses up to f/8.

Other new items on the Nikon D4 include a 3.2-inch LCD with VGA resolution, the ability to enlarge images up to 46x, and an ambient light sensor to automatically brighten or dim the screen based on lighting conditions.

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